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Natural selection can only act on traits that have a narrow sense heritability higher than 0. Why might this be the case, in the context of the breeders equation?
What is: Natural selection can only occur when a trait has a fitness advantage (which means the breeders will have a mean value different than the population average), and it only matters if it can be passed to the next generation. Dominance interactions cannot be transmitted, and neither can environmental effects.